A new early-diverging probainognathian cynodont and a revision of the occurrence of cf. Aleodon from the Chañares Formation, northwestern Argentina: New clues on the faunistic composition of the latest Middle-?earliest Late Triassic Tarjadia Assemblage Zone.

التفاصيل البيبلوغرافية
العنوان:	A new early-diverging probainognathian cynodont and a revision of the occurrence of cf. Aleodon from the Chañares Formation, northwestern Argentina: New clues on the faunistic composition of the latest Middle-?earliest Late Triassic Tarjadia Assemblage Zone.
المؤلفون:	Martinelli AG; Sección Paleontología de Vertebrados, CONICET-Museo Argentino de Ciencias Naturales 'Bernardino Rivadavia', Buenos Aires, Argentina.; Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Buenos Aires, Argentina., Ezcurra MD; Sección Paleontología de Vertebrados, CONICET-Museo Argentino de Ciencias Naturales 'Bernardino Rivadavia', Buenos Aires, Argentina.; Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Buenos Aires, Argentina., Fiorelli LE; Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Buenos Aires, Argentina.; Centro Regional de Investigaciones Científicas y Transferencia Tecnológica de La Rioja (Prov. de La Rioja-UNLaR-SEGEMAR-UNCa-CONICET), La Rioja, Argentina., Escobar J; Sección Paleontología de Vertebrados, CONICET-Museo Argentino de Ciencias Naturales 'Bernardino Rivadavia', Buenos Aires, Argentina.; Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Buenos Aires, Argentina., Hechenleitner EM; Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Buenos Aires, Argentina.; Centro Regional de Investigaciones Científicas y Transferencia Tecnológica de La Rioja (Prov. de La Rioja-UNLaR-SEGEMAR-UNCa-CONICET), La Rioja, Argentina., von Baczko MB; Sección Paleontología de Vertebrados, CONICET-Museo Argentino de Ciencias Naturales 'Bernardino Rivadavia', Buenos Aires, Argentina.; Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Buenos Aires, Argentina., Taborda JRA; Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Buenos Aires, Argentina.; CICTERRA, Ciudad Universitaria, Córdoba, Argentina., Desojo JB; Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Buenos Aires, Argentina.; División Paleontología Vertebrados, Museo de La Plata, Facultad de Ciencias Naturales y Museo, Buenos Aires, Argentina.
المصدر:	Anatomical record (Hoboken, N.J. : 2007) [Anat Rec (Hoboken)] 2024 Apr; Vol. 307 (4), pp. 818-850. Date of Electronic Publication: 2024 Jan 29.
نوع المنشور:	Journal Article
اللغة:	English
بيانات الدورية:	Publisher: John Wiley & Sons Country of Publication: United States NLM ID: 101292775 Publication Model: Print-Electronic Cited Medium: Internet ISSN: 1932-8494 (Electronic) Linking ISSN: 19328486 NLM ISO Abbreviation: Anat Rec (Hoboken) Subsets: MEDLINE
أسماء مطبوعة:	Original Publication: Hoboken, NJ : John Wiley & Sons, 2007-
مواضيع طبية MeSH:	Fossils*, Phylogeny ; Argentina
مستخلص:	The Chañares Formation (Ischigualasto-Villa Unión Basin) is worldwide known by its exquisitely preserved fossil record of latest Middle-to-early Late Triassic tetrapods, including erpetosuchids, "rauisuchians," proterochampsids, gracilisuchids, dinosauromorphs, pterosauromorphs, kannemeyeriiform dicynodonts, and traversodontid, chiniquodontid and probainognathid cynodonts, coming from the Tarjadia (bottom) and Massetognathus-Chanaresuchus (top) Assemblage Zones of its lower member. Regarding cynodonts, most of its profuse knowledge comes from the traditional layers discovered by Alfred Romer and his team in the 1960s that are now enclosed in the Massetognathus-Chanaresuchus Assemblage Zone (AZ). In this contribution we focus our study on the probainognathian cynodonts discovered in levels of the Tarjadia Assemblage Zone. We describe a new chiniquodontid cynodont with transversely broad postcanine teeth (Riojanodon nenoi gen. et sp. nov.) which is related to the genus Aleodon. In addition, the specimen CRILAR-Pv 567 previously referred to cf. Aleodon is here described, compared, and included in a phylogenetic analysis. It is considered as an indeterminate Aleodontinae nov., a clade here proposed to included chiniquodontids with transversely broad upper and lower postcanines, by having a cuspidated sectorial labial margin and a lingual platform that is twice broader than a lingual cingulum. Cromptodon mamiferoides, from the Cerro de Las Cabras Formation (Cuyo Basin), was also included in the phylogenetic analysis and recovered as an Aleodontinae. The new cynodont and the record of Aleodontinae indet. reinforce the faunal differentiation between the Tarjadia and Massetognathus-Chanaresuchus Assemblage Zones, in the lower member of the Chañares Formation, and inform on the diverse chiniquodontid clade with both sectorial and transversely broad postcanine teeth. (© 2024 American Association for Anatomy.)
References:	Abdala, F. (2007). Redescription of Platycraniellus elegans (Therapsida, Cynodontia) from the Lower Triassic of South Africa, and the cladistic relationships of eutheriodonts. Palaeontology, 50, 591-618. Abdala, F. (2021). Permo-Jurassic cynodonts: The early road to mammalness. In D. Alderton & S. A. Ellias (Eds.), Encyclopedia of geology (2nd ed., Vol. 3, pp. 206-226). Academic Press. Abdala, F., Barberena, M. A., & Dornelles, J. (2002). A new species of the traversodontid cynodont Exaeretodon from the Santa Maria Formation (Middle/Late Triassic) of southern Brazil. Journal of Vertebrate Paleontology, 22, 313-325. Abdala, F., Gaetano, L. C., Martinelli, A. G., Soares, M. B., Hancox, P. J., & Rubidge, B. S. (2020). Non-mammaliaform cynodonts from western Gondwana and the significance of Argentinean forms in enhancing understanding of the group. Journal of South American Earth Sciences, 104, 102884. https://doi.org/10.1016/j.jsames.2020.102884. Abdala, F., Gaetano, L. C., Smith, R. M. H., & Bruce, R. (2019). A new large cynodont from the late Permian (Lopingian) of the South African Karoo Basin and its phylogenetic significance. Zoological Journal of the Linnean Society, 186, 983-1005. Abdala, F., & Giannini, N. P. (2000). Gomphodont cynodonts of the Chañares Formation: The analysis of an ontogenetic sequence. Journal of Vertebrate Paleontology, 20, 501-506. Abdala, F., & Giannini, N. P. (2002). Chiniquodontid cynodonts: Systematic and morphometric considerations. Palaeontology, 45, 1151-1170. Abdala, F., Jasinoski, S. C., & Fernandez, V. (2013). Ontogeny of the Early Triassic cynodont Thrinaxodon liorhinus (Therapsida): Dental morphology and replacement. Journal of Vertebrate Paleontology, 33(6), 1408-1431. Abdala, F., Marsicano, C. A., Smith, R. M. H., & Swart, R. (2013). Strengthening Western Gondwanan correlations: A Brazilian dicynodont (Synapsida, Anomodontia) in the Middle Triassic of Namibia. Gondwana Research, 23(3), 1151-1162. Abdala, F., Martinelli, A. G., & González, F. R. (2022). La paleoherpetología en el Instituto Miguel Lillo de la Universidad Nacional de Tucumán, Argentina. Publicación Electrónica de la Asociación Paleontológica Argentina, 22(1), 67-87. https://doi.org/10.5710/PEAPA.16.05.2021.360. Abdala, F., Neveling, J., & Welman, J. (2006). A new trirachodontid cynodont from the lower levels of the Burgersdorp formation (Lower Triassic) of the Beaufort group, South Africa and the cladistic relationships of Gondwanan gomphodonts. Zoological Journal of the Linnean Society, 147, 383-413. Abdala, F., & Ribeiro, A. M. (2003). A new traversodontid cynodont from the Santa Maria Formation (Ladinian-Carnian) of southern Brazil, with a phylogenetic analysis of Gondwanan traversodontids. Zoological Journal of the Linnean Society, 139, 529-545. Abdala, F., & Smith, R. M. H. (2009). A Middle Triassic cynodont fauna from Namibia and its implications for the biogeography of Gondwana. Journal of Vertebrate Paleontology, 29, 837-851. Agnolin, F. L., & Ezcurra, M. D. (2019). The validity of Lagosuchus talampayensis Romer, 1971 (Archosauria, Dinosauriformes): Implications for early dinosauriform systematics. Breviora, 565, 1-21. Agnolin, F., Brissón Egli, F., Ezcurra, M. D., Langer, M. C. and Novas, F. (2022). New specimens provide insights into the anatomy of the dinosauriform Lewisuchus admixtus Romer, 1972 from the upper Triassic levels of the Chañares Formation, NW Argentina. The Anatomical Record 305(5), 1119-1146. https://doi.org/10.1002/ar.24731. Angielczyk, K. D., Hancox, P. J., & Nabavizadeh, A. (2018). A redescription of the Triassic kannemeyeriiform dicynodont Sangusaurus (Therapsida, Anomodontia), with an analysis of its feeding system. In: C. A. Sidor and S. J. Nesbitt (eds.), Vertebrate and climatic evolution in the Triassic rift basins of Tanzania and Zambia. Society of Vertebrate Paleontology Memoir 17. Journal of Vertebrate Paleontology, 37(6, Supplement), 189-227. Angielczyk, K. D., & Kammerer, C. F. (2018). Non-mammalian synapsids: The deep roots of the mammalian family tree. In F. Zachos & R. Asher (Eds.), Mammalian evolution, diversity and systematics. Handbook of zoology (pp. 117-198). De Gruyter. Ávila, J. N., Chemale, F., Jr., Mallmann, G., Kawashita, K., & Armstrong, R. (2006). Combined stratigraphic and isotopic studies of Triassic strata, Cuyo Basin, Argentine Precordillera. Geological Society of America Bulletin, 118, 1088-1098. https://doi.org/10.1130/B25893.1. Battail, B. (1991). Les cynodontes (Reptilia, Therapsida): Une phylogénie. Bulletin du Museum Nationale d'Histoire Naturelle, 13, 17-105. Benoit, J., Nxumalo, M., Norton, L. A., Fernandez, V., Gaetano, L. C., Rubidge, B., & Abdala, F. (2022). Synchrotron scanning sheds new light on Lumkuia fuzzi (Therapsida, Cynodontia) from the Middle Triassic of South Africa and its phylogenetic placement. Journal of African Earth Sciences, 196, 104689. Benoit, J., Ruf, I., Miyamae, J. A., Fernandez, V., Rodrigues, P. G., & Rubidge, B. S. (2020). The evolution of the maxillary canal in Probainognathia (Cynodontia, Synapsida): Reassessment of the homology of the infraorbital foramen in mammalian ancestors. Journal of Mammalian Evolution, 27, 329-348. Bonaparte, J. F. (1962). Descripción del cráneo y mandíbula de Exaeretodon frenguellii, Cabrera y su comparación con Diademodontidae, Tritylodontidae y los cinodontes sudamericanos. Publicaciones del Museo Municipal de Ciencias Naturales y Tradicional de Mar del Plata, 1, 135-202. Bonaparte, J. F. (1963a). Un nuevo cinodonte gonfodonte del Triásico Medio Superior de San Juan, Proexaeretodon vincei n. gen., n. sp. Acta Geológica Lilloana, 4, 129-133. Bonaparte, J. F. (1963b). Descripción del esqueleto postcraneano de Exaeretodon. Acta Geológica Lilloana, 4, 5-53. Bonaparte, J. F. (1963c). Descripción de Ischignathus sudamericanus n. gen. N. sp., nuevo cinodonte gonfodonte del Triásico Medio Superior de San Juan, Argentina. Acta Geológica Lilloana, 5, 111-128. Bonaparte, J. F. (1966). Chiniquodon Huene (Therapsida-Cynodontia) en el Triásico de Ischigualasto, Argentina. Acta Geologica Lilloana, 8, 157-169. Bonaparte, J. F. (1969). Dos nuevas “faunas” de reptiles Triásicos de Argentina. In 1st International IUGS Symposium of Gondwana (pp. 283-306). UNESCO. Bonaparte, J. F. (1971). Los tetrápodos del sector superior de la Formación Los Colorados, La Rioja, Argentina (Triásico Superior). Op Lilloana, 22, 1-183. Bonaparte, J. F. (1972). Cromptodon mamiferoides, Galesauridae de la Formación Río Mendoza, Mendoza, Argentina (Therapsida-Cynodontia). Ameghiniana, 9, 343-353. Bonaparte, J. F. (1975). Nuevos materiales de Lagosuchus talampayensis Romer y su significado en el origen de los Saurischia. Acta Geológica Lilloana, 13, 1-90. Bonaparte, J. F. (1981). Nota sobre una nueva fauna del Triásico Inferior del Sur de Mendoza, Argentina, correspondiente a la Zona de Lystrosaurus (Dicynodontia-Proterosuchia). Anais Congreso Latino-Americano de Paleontologia, Abstract book, 277-288. Bonaparte, J. F. (1997). El Triásico de San Juan - La Rioja, Argentina, y sus dinosaurios (p. 190). Museo Argentino Ciencias Naturales. Bonaparte, J. F., Martinelli, A. G., & Schultz, C. L. (2005). New information on Brasilodon and Brasilitherium (Cynodontia, Probainognathia) from the Late Triassic of southern Brazil. Revista Brasileira de Paleontologia, 8, 25-46. Bonaparte, J. F., Martinelli, A. G., Schultz, C. L., & Rubert, R. (2003). The sister group of mammals: Small cynodonts from the Late Triassic of southern Brazil. Revista Brasileira de Paleontologia, 5, 5-27. Bonaparte, J. F., & Migale, L. A. (2015). Protomamíferos y Mamíferos Mesozoicos de América del Sur. Mercedes, Buenos Aires, Argentina (2st ed.). Mercedes. Brink, A. S. (1977). A model of tooth replacement in the “mammal-like reptile” Diademodon. South African Journal of Science, 73, 138-143. Broili, F., & Schröder, J. (1934). Zur osteologie des kopfes von Cynognathus. Sitzugsberichte der Bayerischen, Akademie der Wissenschaften, 1934, 95-128. Broom, R. (1905). On the use of the term Anomodontia. Record of the Albany Museum, 1, 266-269. Cabrera, A. (1943). El primer hallazgo de terápsidos en la Argentina. Notas del Museo de La Plata, Paleontología, 8, 317-331. Casamiquela, R. M. (1961). Dos nuevos estagonolepoideos argentinos (de Ischigualasto, San Juan). Revista de la Asociación Geológica Argentina, 16(3-4), 143-204. Casamiquela, R. M. (1967). Un nuevo dinosaurio ornitisquio Triásico (Pisanosaurus mertii; Ornithopoda) de la Formación Ischigualasto, Argentina. Ameghiniana, 4, 47-64. Cohen, K. M., Finney, S. C., Gibbard, P. L., & Fan, J.-X. (2013). The ICS international chronostratigraphic chart. Episodes, 36, 199-204. Crompton, A. W. (1955). On some Triassic cynodonts from Tanganyika. Proceedings. Zoological Society of London, 1955(125), 617-669. Crompton, A. W. (1972). Postcanine occlusion in cynodonts and tritylodontids. Bulletin of the British Museum (Natural History)., 21(2), 27-71. Crompton, A. W., Musinsky, C., & Owerkowicz, T. (2015). Evolution of the mammalian nose. In K. Dial, N. Shubin, & E. Brainerd (Eds.), Great transformations in vertebrate evolution (pp. 189-203). University of Chicago Press. Crompton, A. W., Owerkowivcz, T., Bhullar, B. A. S., & Musinsky, C. (2017). Structure of the nasal region of non-mammalian cynodonts and mammaliaforms: Speculations on the evolution of mammalian endothermy. Journal of Vertebrate Paleontology, 37, e1269116. Cromton, A. W. (1974). The dentitions and relationships of the southern African Triassic mammals, Erythrotherium parringtoni and Megazostrodon rudnerae. Bulletin of the British Museum (Natural History). Geology, 24, 397-437. Cruickshank, A. R. I. (1967). A new dicynodont genus from the Manda Formation of Tanzania (Tanganyika). Journal of Zoology, 153(2), 163-208. https://doi.org/10.1111/j.1469-7998.1967.tb04059.x. Desojo, J. B., & Arcucci, A. B. (2009). New material of Luperosuchus fractus (Archosauria: Crurotarsi) from the Middle Triassic of Argentina: The earliest known South American ‘rauisuchian’. Journal of Vertebrate Paleontology, 29(4), 1311-1315. Desojo, J. B., & Báez, A. M. (2007). Cranial morphology of the Late Triassic South American archosaur Neoaetosauroides engaeus: Evidence for aetosaurian diversity. Palaeontology, 50, 267-276. Desojo, J. B., Fiorelli, L. E., Ezcurra, M. D., Martinelli, A. G., Ramezani, J., Da Rosa, Á. A. S., von Baczko, M. B., Trotteyn, M. J., Montefeltro, F. C., Ezpeleta, M., & Langer, M. C. (2020). The Late Triassic Ischigualasto Formation at Cerro Las Lajas (La Rioja, Argentina): Fossil tetrapods, high-resolution chronostratigraphy, and faunal correlations. Scientific Reports, 10, 12782. https://doi.org/10.1038/s41598-020-67854-1. Domnanovich, N. S., & Marsicano, C. A. (2012). The Triassic dicynodont Vinceria (Therapsida, Anomodontia) from Argentina and a discussion on basal Kannemeyeriiformes. Geobios, 45(2), 173-186. Escobar, J., Martinelli, A., Ezcurra, M., Fiorelli, L., von Baczko, M. B., Novas, F., & Desojo, J. (2023). A reassessment of the mandibular anatomy of Dinodontosaurus brevirostris (Synapsida, Dicynodontia) from the Ladinian-early Carnian Chañares Formation (northwestern Argentina), and its taxonomic and phylogenetic significance. Ameghiniana, 60(2), 178-201. Escobar, J. A., Martinelli, A. G., Ezcurra, M. D., Fiorelli, L. E., & Desojo, J. B. (2021). A new stahleckeriid dicynodont record from the late Ladinian-? early Carnian levels of the Chañares Formation (Ischigualasto-Villa Unión Basin) of northwestern Argentina. Journal of South American Earth Sciences, 109, 103275. Ezcurra, M. D. (2023). Exploring different weightings against homoplasy in genealogies of paleontological phylogenetic matrices. In Abstract book annual meeting of the Willi Hennig society XL: 8. Cornell University. Ezcurra, M. D., Bandyopadhyay, S., & Gower, D. J. (2021). A new erythrosuchid archosauriform from the Middle Triassic Yerrapalli formation of south-central India. Ameghiniana, 58, 132-168. Ezcurra, M. D., & Butler, R. J. (2018). The rise of the ruling reptiles and ecosystem recovery from the Permo-Triassic mass extinction. Proceedings of the Royal Society B, 285(1880), 20180361. Ezcurra, M. D., Fiorelli, L. E., Martinelli, A. G., Rocher, S. M., Baczko, M. B., Ezpeleta, M., Taborda, J. R. A., Hechenleitner, E. M., Trotteyn, M. J., & Desojo, J. B. (2017). Deep faunistic changes preceded the raise of dinosaurs in southwestern Pangaea. Nature Ecology and Evolution, 1, 1477-1483. Ezcurra, M. D., Fiorelli, L. E., Trotteyn, M. J., Martinelli, A. G., & Desojo, J. B. (2021). The rhynchosaur record, including a new stenaulorhynchine taxon, from the Chañares Formation (upper Ladinian-? lowermost Carnian levels) of La Rioja Province, north-western Argentina. Journal of Systematic Palaeontology, 18, 1907-1938. https://doi.org/10.1080/14772019.2020.1856205. Ezcurra, M. D., Montefeltro, F. C., Pinheiro, F. L., Trotteyn, M. J., Gentil, A. R., Lehmann, O. E., & Pradelli, L. A. (2021). The stem-archosaur evolutionary radiation in South America. Journal of South American Earth Sciences, 105, 102935. https://doi.org/10.1016/j.jsames.2020.102935. Ezcurra, M. D., Nesbitt, S. J., Bronzati, M., Dalla Vecchia, F. B., Agnolin, F. L., Benson, R. B. J., Brissón Egli, F., Cabreira, S. F., Evers, S. W., Gentil, A. R., Irmis, R. B., Martinelli, A. G., Novas, F. E., Roberto da Silva, L., Smith, N. D., Stocker, M. R., Turner, A. H., & Langer, M. C. (2020). Enigmatic dinosaur precursors bridge the gap to the origin of Pterosauria. Nature, 588, 445-449. https://doi.org/10.1038/s41586-020-3011-4. Ezcurra, M. D., Nesbitt, S. J., Fiorelli, L. E., & Desojo, J. B. (2020). New specimen sheds light on the anatomy and taxonomy of the early Late Triassic dinosauriforms from the Chañares Formation, NW Argentina. The Anatomical Record, 303, 1393-1438. Ezcurra, M. D., Baczko, M. B., Trotteyn, M. J., & Desojo, J. B. (2019). New proterochampsid specimens expand the morphological diversity of the rhadinosuchines of the Chañares Formation (lower Carnian, northwestern Argentina). Ameghiniana, 56, 79-115. Ezcurra, M. D., Trotteyn, M. J., Fiorelli, L. E., von Baczko, M. B., Taborda, J. R. A., Iberlucea, M., & Desojo, J. B. (2014). The oldest rhynchosaur from Argentina: A Middle Triassic rhynchosaurid from the Chañares Formation (Ischigualasto-Villa Unión Basin, La Rioja Province). Paläontologische Zeitschrift, 88, 453-460. https://doi.org/10.1007/s12542-013-0203-3. Fedorov, A., Beichel, R., Kalpathy-Cramer, J., Finet, J., Fillion-Robin, J.-C., Pujol, S., Bauer, C., Jennings, D., Fennessy, F. M., Sonka, M., Buatti, J., Aylward, S. R., Miller, J. V., Pieper, S., & Kikinis, R. (2012). 3D slicer as an image computing platform for the quantitative imaging network. Magnetic Resonance Imaging., 30(9), 1323-1341. https://doi.org/10.1016/j.mri.2012.05.001. Fiorelli, L. E., Ezcurra, M. D., Hechenleitner, E. M., Argañaraz, E., Taborda, J. R. A., Trotteyn, M. J., Baczko, M. B., & Desojo, J. B. (2013). The oldest known communal latrines provide evidence of gregarism in Triassic megaherbivores. Scientific Reports, 3, 3348. Fiorelli, L. E., Rocher, S., Martinelli, A. G., Ezcurra, M. D., Hechenleitner, E. M., & Ezpeleta, M. (2018). Tetrapod burrows from the Middle-Upper Triassic Chañares Formation (La Rioja, Argentina) and its palaeoecological implications. Palaeogeography, Palaeoclimatology, Palaeoecology, 496, 85-102. Foth, C., Ezcurra, M. D., Sookias, R., Brusatte, S. L., & Butler, R. J. (2016). Unappreciated diversification of stem archosaurs during the Middle Triassic predated the dominance of dinosaurs. BMC Evolutionary Biology, 16(1), 188. https://doi.org/10.1186/s12862-016-0761-6. Foth, C., Sookias, R. B., & Ezcurra, M. D. (2021). Rapid initial morphospace expansion and delayed morphological disparity peak in the first 100 million years of the archosauromorph evolutionary radiation. Frontiers in Earth Science, 9, 723973. https://doi.org/10.3389/feart.2021.723973. Franco, A. S., Müller, R. T., Martinelli, A. G., Hoffmann, C. A., & Kerber, L. (2021). The nasal cavity of two traversodontid cynodonts (Eucynodontia, Gomphodontia) from the Upper Triassic of Brazil. Journal of Paleontology, 95(4), 845-860. Gaetano, L. C., Abdala, F., Seoane, F. D., Tartaglione, A., Schulz, M., Otero, A., Leardi, J. M., Apaldetti, C., Krapovickas, V., & Steimbach, E. (2022). A new cynodont from the Upper Triassic Los Colorados Formation (Argentina, South America) reveals a novel paleobiogeographic context for mammalian ancestors. Scientific Reports, 12, 6451. https://doi.org/10.1038/s41598-022-10486-4. Goloboff, P. A., Farris, J. S., Källersjö, M., Oxelman, B., Ramírez, M. J., & Szumik, C. A. (2003). Improvements to resampling measures of group support. Cladistics, 19(4), 324-332. Goloboff, P. A., & Morales, M. E. (2023). TNT version 1.6, with a graphical interface for MacOS and Linux, including new routines in parallel. Cladistics, 39(2), 144-153. Goñi, R., & Goin, F. J. (1987). El origen de los postcaninos gonfodontes de Andescynodon mendozensis Bonaparte (Cynodontia, Traversodontidae). Ameghiniana, 24(3-4), 235-239. Gouiric-Cavalli, S., Desojo, J. B., Ezcurra, M. D., Fiorelli, L. E., & Martinelli, A. G. (2017). First fish remains from the early Late Triassic Chañares Formation (La Rioja, Argentina) and their paleobiogeographic implications. Ameghiniana, 54(2), 137-150. https://doi.org/10.5710/AMGH.10.10.2016.3015. Gow, C. E. (1980). The dentitions of the Tritheledontidae (Therapsida: Cynodontia). Proceedings of the Royal Society of London, Series B, 208, 461-481. Grine, F. E. (1977). Postcanine tooth function and jaw movement in the gomphodont cynodont Diademodon (Reptilia; Therapsida). Palaeontologia Africana, 20, 123-135. Guignard, M. L., Martinelli, A. G., & Soares, M. B. (2019a). Postcranial anatomy of Riograndia guaibensis (Cynodontia: Ictidosauria). Geobios, 53, 9-21. Guignard, M. L., Martinelli, A. G., & Soares, M. B. (2019b). The postcranial anatomy of Brasilodon quadrangularis and the acquisition of mammaliaform traits among non-mammaliaform cynodonts. PLoS One, 14(5), e0216672. Gulbranson, E. L., Ciccioli, P. L., Montañez, I. P., Marenssi, S. A., Limarino, C. O., Schmitz, M. D., & Davydov, V. (2015). Paleoenvironments and age of the Talampaya Formation: The Permo-Triassic boundary in northwestern Argentina. Journal of South American Earth Sciences, 63, 310-322. https://doi.org/10.1016/j.jsames.2015.08.008. Hancox, P. J., Angielczyk, K. D., & Rubidge, B. S. (2013). Angonisaurus and Shansiodon, dicynodonts (Therapsida, Anomodontia) from Subzone C of the Cynognathus Assemblage Zone (Middle Triassic) of South Africa. Journal of Vertebrate Paleontology, 33(3), 655-676. Hechenleitner, E. M., Martinelli, A. G., Juarez, M., Fiorelli, L. E., Rocher, S., Taborda, J. R. A., Leuzinger, L., de la Vega, S. R., Acosta, L., & Desojo, J. B. (2023). Una nueva asociación de vertebrados continentales del Triásico tardío, Formación Santo Domingo, Quebrada Santo Domingo, La Rioja, Argentina. In Libro de Resúmenes, 36 Jornadas Argentinas de Paleontología de Vertebrados, La Rioja, Argentina. CRILAR. Hendrickx, C., Gaetano, L. C., Choiniere, J. N., Mocke, H., & Abdala, F. (2020). A new traversodontid cynodont with a peculiar postcanine dentition from the Middle/Late Triassic of Namibia and dental evolution in basal gomphodonts. Journal of Systematic Palaeontology, 18(20), 1669-1706. Hillenius, W. J. (1992). The evolution of nasal turbinates and mammalian endothermy. Paleobiology, 18, 17-29. Hillenius, W. J. (1994). Turbinates in therapsids: Evidence for late Permian origins of mammalian endothermy. Evolution, 48, 207-229. Hoffman, C., de Andrade, M., & Martinelli, A. G. (2023). Anatomy of the holotype of “Probelesodon kitchingi” revisited, a chiniquodontid cynodont (Synapsida, Probainognathia) from the early Late Triassic of southern Brazil. Journal of Paleontology, 97(3), 693-710. https://doi.org/10.1017/jpa.2023.25. Hopson, J. A. (1990). Cladistic analysis of therapsid relationships. Journal of Vertebrate Paleontology, 10(3 Supplement), 28A. Hopson, J. A. (1991). Systematics of the nonmammalian Synapsida and implications for patterns of evolution in synapsids. In H. Schultze & L. Trueb (Eds.), Origins of the higher groups of Tetrapods: Controversy and consensus (pp. 635-694). Cornell University Press. https://doi.org/10.7591/9781501718335-020. Hopson, J. A., & Kitching, J. W. (2001). A probainognathian cynodont from South Africa and the phylogeny of nonmammalian cynodonts. Bulletin of the Museum of Comparative Zoology, 156, 5-35. Huttenlocker, A. K., & Sidor, C. A. (2020). A basal nonmammaliaform cynodont from the Permian of Zambia and the origins of mammalian endocranial and postcranial anatomy. Journal of Vertebrate Paleontology, 40(5), e1827413. https://doi.org/10.1080/02724634.2020.1827413. Jasinoski, S. C., & Chinsamy, A. (2012). Mandibular histology and growth of the non-mammaliaform cynodont Tritylodon. Journal of Anatomy, 220, 564-579. Jenkins, F. A., Jr. (1970). The Chañares (Argentina) Triassic reptile fauna VII. The postcranial skeleton of the traversodontid Massetognathus pascuali (Therapsida, Cynodontia). Breviora, 352, 1-28. Jensen, J. A. (2001). The road to Chilecito. Queen Victoria Museum and Art Gallery. Kalandadze, N. N. (1970). New Triassic kanemeyeriids from the South Ural region. In K. K. Flerov (Ed.), Materials on evolution of terrestrial vertebrates (pp. 51-57). Nauka Press [in Russian]. Kammerer, C., Flynn, J. J., Ranivoharimanana, L., & Wyss, A. R. (2010). The first record of a probainognathian (Cynodontia: Chiniquodontidae) from the Triassic of Madagascar. Journal of Vertebrate Paleontology, 30(6), 1889-1894. Kammerer, C. F. (2016). A new taxon of cynodont from the Tropidostoma Assemblage Zone (upper Permian) of South Africa, and the early evolution of Cynodontia. Papers in Palaeontology, 2(3), 387-397. Kammerer, C. F., Angielczyk, K. D., & Nesbitt, S. J. (2018). Novel hind limb morphology in a kannemeyeriiform dicynodont from the Manda beds (Songea group, Ruhuhu Basin) of Tanzania. In: Sidor, C.A., Nesbitt, S.J. (Eds.), Vertebrate and climatic evolution in the Triassic rift basins of Tanzania and Zambia. Society of Vertebrate Paleontology Memoir 17. Journal of Vertebrate Paleontology, 37(6, supplement), 178-188. Kammerer, C. F., & Ordoñez, M. A. (2021). Dicynodonts (Therapsida: Anomodontia) of South America. Journal of South American Earth Sciences, 108, 103171. https://doi.org/10.1016/j.jsames.2021.103171. Kemp, T. S. (1980). Aspects of the structure and functional anatomy of the Middle Triassic cynodont Luangwa. Journal of Zoology, 191, 193-239. Kemp, T. S. (1982). Mammal-like reptiles and the origin of mammals (pp. 1-363). Academic Press. Kemp, T. S. (2005). The origin and evolution of mammals (p. 342). Oxford University Press on Demand. Kerber, L., Martinelli, A. G., Müller, R. T., & Pretto, F. A. (2022). A new specimen provides insights into the anatomy of Irajatherium hernandezi, a poorly known probainognathian cynodont from the Late Triassic of southern Brazil. The Anatomical Record, 305(11), 3113-3132. Kerber, L., Martinelli, A. G., Rodrigues, P. G., Ribeiro, A. M., Schultz, C. L., & Soares, M. B. (2020). New record of Prozostrodon brasiliensis (Eucynodontia: Prozostrodontia) from its type-locality (Upper Triassic, southern Brazil): Comments on the endocranial morphology. Revista Brasileira de Paleontologia, 23, 259-269. Kerber, L., Miron, L. R., Budabué, J. M., & Martinelli, A. G. (2024). Endocranial anatomy of the early prozostrodonts (Eucynodontia: Probainognathia) and the neurosensory evolution of the mammalian forerunners. The Anatomical Record, 307(4), 1442-1473. https://doi.org/10.1002/ar.25215. Kerber, L., Pretto, F. A., & Müller, R. T. (2024). New information on the mandibular anatomy of Agudotherium gassenae, a Late Triassic non-mammaliaform probainognathian from Brazil. The Anatomical Record, 307(4), 1515-1523. https://doi.org/10.1002/ar.25317. Kermack, K. A., Mussett, F., & Rigney, H. W. (1981). The skull of Morganucodon. Zoological Journal of the Linnean Society, 71, 1-158. Kielan-Jaworowska, Z., Cifelli, R. L., & Luo, Z.-X. (2004). Mammals from the age of dinosaurs: Origins, evolution, and structure. Columbia University Press. Kokogián, D. A., Spalletti, L., Morel, E., Artabe, A., Martínez, R. N., Alcober, O. A., Milana, J. P., Zavattieri, A. M., & Papú, O. H. (1999). Los depósitos continentales triásicos. Instituto de Geología y Recursos Minerales, Geología Argentina, Anales, 29(15), 377-398. Langer, M. C., Ribeiro, A. A. M., Schultz, C. L., & Ferigolo, J. (2007). The continental tetrapod-bearing Triassic of south Brazil. Bulletin of the American Museum of Natural History, 41, 201-218. Lecuona, A., Desojo, J. B., & Cerda, I. A. (2020). New information on the anatomy and histology of Gracilisuchus stipanicicorum (Archosauria: Pseudosuchia) from the Chañares Formation (early Carnian), Argentina. Comptes Rendus Palevol, 19(3), 40-62. Liu, J. (2022). On kannemeyeriiform dicynodonts from the Shaanbeikannemeyeria Assemblage Zone of the Ordos Basin, China. Vertebrata Palasiatica, 60(3), 212-248. https://doi.org/10.19615/j.cnki.2096-9899.220601. Liu, J. & Olsen, P. (2010). The phylogenetic relationships of Eucynodontia (Amniota: Synapsida). Journal of Mammalian Evolution 17, 151-176. Liu, J., & Abdala, F. (2014). Phylogeny and taxonomy of the Traversodontidae. In C. F. Kammerer, K. D. Angielczyk, & J. Fröbisch (Eds.), Early evolutionary history of the Synapsida (pp. 255-279). Springer. Liu, J., & Powell, J. (2009). Osteology of Andescynodon (Cynodontia, Traversodontidae) from the Middle Triassic of Argentina. American Museum Novitates, 3674, 1-19. Lukic-Walther, M., Brocklehurst, N., Kammerer, C., & Fröbisch, J. (2019). Diversity patterns of nonmammalian cynodonts (Synapsida, Therapsida) and the impact of taxonomic practice and research history on diversity estimates. Paleobiology, 45(1), 56-69. https://doi.org/10.1017/pab.2018.38. Luo, Z.-X. (1994). Sister-group relationships of mammals and transformations of diagnostic mammalian characters. In N. C. Fraser & H.-D. Sues (Eds.), The shadow of the dinosaurs: Early mesozoic tetrapods (pp. 98-128). Cambridge, Cambridge University Press. Mancuso, A. C., Gaetano, L. C., Leardi, J. M., Abdala, F., & Arcucci, A. B. (2014). The Chañares Formation: A window to a Middle Triassic tetrapod community. Lethaia, 47(2), 244-265. https://doi.org/10.1111/let.12055. Marsicano, C. A., Irmis, R. B., Mancuso, A. C., Mundil, R., & Chemale, F. (2015). The precise temporal calibration of dinosaur origins. Proceedings of the National Academy of Sciences of the United States of America, 113, 509-513. Martinelli, A. G. (2010). On the postcanine dentition of Pascualgnathus polanskii Bonaparte (Cynodontia: Traversodontidae) from the Middle Triassic of Argentina. Geobios, 43, 629-638. Martinelli, A. G. (2017). Contribuição ao conhecimento dos cinodontes probainognátios (Therapsida, Cynodontia, Probainognathia) do Triássico da América do Sul e seu impacto na origem dos Mammaliaformes. PhD Thesis, Universidade Federal do Rio Grande do Sul, Brazil. Martinelli, A. G., Agnolín, F. L., Ezcurra, M. D., Isasi, M. P., & Novas, F. E. (2022). El Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” y los aportes a la paleoherpetología Argentina. Publicación Electrónica de la Asociación Paleontológica Argentina, 22(1), 188-2007. https://doi.org/10.5710/PEAPA.14.05.2021.356. Martinelli, A. G., & Bonaparte, J. F. (2011). Postcanine replacement in Brasilodon and Brasilitherium (Cynodontia, Probainognathia) and its bearing in cynodont evolution. In J. Calvo, J. Porfiri, B. González Riga, & D. Dos Santos (Eds.), Dinosaurios y Paleontología desde América Latina. Anales del III Congreso Latinoamericano de Paleontología (pp. 179-186). Editorial de la Universidad Nacional de Cuyo. Martinelli, A. G., Bonaparte, J. F., Schultz, C. L., & Rubert, R. (2005). A new tritheledontid (Therapsida, Eucynodontia) from the Late Triassic of Rio Grande do Sul (Brazil) and its phylogenetic relationships among carnivorous non-mammalian eucynodonts. Ameghiniana, 42, 191-208. Martinelli, A. G., de la Fuente, M., & Abdala, F. (2009). Diademodon tetragonus Seeley, 1894 (Therapsida: Cynodontia) in the Triassic of South America and its biostratigraphic implications. Journal of Vertebrate Paleontology, 29, 852-862. Martinelli, A. G., Eltink, E., Da-Rosa, Á. A. S., & Langer, M. C. (2017). A new cynodont (Therapsida) from the Hyperodapedon Assemblage Zone (upper Carnian-Norian) of southern Brazil improves the Late Triassic probainognathian diversity. Papers in Palaeontology, 3, 401-423. Martinelli, A. G., Ezcurra, M. D., Fiorelli, L. E., Escobar, J. A., von Baczko, M. B., Hechenleitner, M., Trotteyn, J., & Desojo, J. B. (2023). The cynodont Menadon (Traversodontidae) in the early Late Triassic of the Ischigualasto-Villa Unión Basin, NW Argentina: First record of a possible new faunal association in the upper Chañares Formation and its gondwanan biostratigraphic implications. In Libro de Resúmenes, 36 Jornadas Argentinas de Paleontología de Vertebrados, La Rioja, Argentina. CRILAR. Martinelli, A. G., Kammerer, C. F., Melo, T. P., Paes Neto, V. D., Ribeiro, A. M., Da-Rosa, A. A. S., Schultz, C. L., & Soares, M. B. (2017). The African cynodont Aleodon (Cynodontia, Probainognathia) in the Triassic of southern Brazil and its biostratigraphic significance. PLoS One, 12(6), e0177948. https://doi.org/10.1371/journal.pone.0177948. Martinelli, A. G., & Rougier, G. W. (2007). On Chaliminia musteloides Bonaparte (Cynodontia, Tritheledontidae) and the phylogeny of the Ictidosauria. Journal of Vertebrate Paleontology, 27, 442-460. Martinelli, A. G., & Soares, M. B. (2016). Evolution of South American cynodonts. Contribuciones del Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, 6, 183-197. Martinelli, A. G., Soares, M. B., de Oliveira, T. V., Rodrigues, P. G., & Schultz, C. L. (2017). The Triassic eucynodont Candelariodon barberenai revisited and the early diversity of stem prozostrodontians. Acta Palaeontologica Polonica, 62, 527-542. Martinelli, A. G., Soares, M. B., & Schwanke, C. (2016). Two new cynodonts (Therapsida) from the Middle-early Late Triassic of Brazil and comments on South American probainognathians. PLoS One, 11, e0162945. Martínez, R. N., Apaldetti, C., Alcober, O. A., Colombi, C. E., Sereno, P. C., Fernandez, E., Santi Malnis, P., Correa, G. A., & Abelin, D. (2013). Vertebrate succession in the Ischigualasto Formation. Soc Vertebr Paleont mem 12. Journal of Vertebrate Paleontology, 32(6, supplement), 10-30. Martínez, R. N., Apaldetti, C., Correa, G. A., Colombi, C. E., Fernández, E., Santi Malnis, P., Praderio, A., Abelín, D., Benegas, L. G., Aguilar-Cameo, A., & Alcober, O. A. (2015). A new Late Triassic vertebrate assemblage from northwestern Argentina. Ameghiniana, 52, 379-390. Martínez, R. N., Fernández, E., & Alcober, O. A. (2013). A new nonmammaliaform eucynodont from the Carnian-Norian Ischigualasto Formation, northwestern Argentina. Revista Brasilera de Paleontología, 16, 61-76. Martínez, R. N., & Forster, C. A. (1996). The skull of Probelesodon sanjuanensis, sp. nov., from the Late Triassic Ischigualasto Formation of Argentina. Journal of Vertebrate Paleontology, 16, 285-291. Martínez, R. N., May, C. L., & Forster, C. A. (1996). A new carnivorous cynodont from the Ischigualasto Formation (Late Triassic, Argentina), with comments on eucynodont phylogeny. Journal of Vertebrate Paleontology, 16, 271-284. Martínez, R. N., Sereno, P. C., Alcober, O. A., Colombi, C. E., Renne, P. R., Montañez, I. P., & Currie, B. S. (2011). A basal dinosaur from the dawn of the dinosaur era in southwestern Pangaea. Science, 331, 206-210. Martínez, R. N., Simões, T. R., Sobral, G., & Apesteguía, S. (2021). A Triassic stem lepidosaur illuminates the origin of lizard-like reptiles. Nature, 597, 235-238. Melo, T. P., Martinelli, A. G., & Soares, M. B. (2017). A new gomphodont cynodont (Traversodontidae) from the Middle-Late Triassic Dinodontosaurus Assemblage Zone of the Santa Maria Supersequence, Brazil. Palaeontology, 60, 571-582. https://doi.org/10.1111/pala.12302. Melo, T. P., Martinelli, A. G., & Soares, M. B. (2022). New occurrences of massetognathine traversodontids and chiniquodontids (Synapsida, Cynodontia) from the early Late Triassic Santacruzodon Assemblage Zone (Santa Maria Supersequence, southern Brazil): Geographic and biostratigraphic implications. Journal of South American Earth Sciences, 115, 103757. https://doi.org/10.1016/j.jsames.2022.103757. Melo, T. P., Ribeiro, A. M., Martinelli, A. G., & Soares, M. B. (2019). Earliest evidence of molariform hypsodonty in a Triassic stem-mammal. Nature Communications, 10, 2841. Mocke, H. B., Gaetano, L. C., & Abdala, F. (2020). A new species of the carnivorous cynodont Chiniquodon (Cynodontia, Chiniquodontidae) from the Namibian Triassic. Journal of Vertebrate Paleontology, 39(6), e1754231. Nesbitt, S., & Desojo, J. B. (2017). The osteology and phylogenetic position of Luperosuchus fractus (Archosauria: Loricata) from the latest Middle Triassic or earliest Late Triassic of Argentina. Ameghiniana, 54(3), 261-282. Norton, L. A., Abdala, F., Rubidge, B. S., & Botha, J. (2020). Tooth replacement patterns in the Early Triassic epicynodont Galesaurus planiceps (Therapsida, Cynodontia). PLoS One, 15, e0243985. https://doi.org/10.1371/journal.pone.0243985. Novas, F. E., Agnolin, F. L., Ezcurra, M. D., Müller, R. T., Martinelli, A. G., & Langer, M. C. (2021). Review of the fossil record of early dinosaurs from South America, and its phylogenetic implications. Journal of South American Earth Sciences, 110, 103341. Oliveira, T. V., Soares, M. B., & Schultz, C. L. (2010). Trucidocynodon riograndensis gen. nov. et sp. nov. (Eucynodontia), a new cynodont from the Brazilian Upper Triassic (Santa Maria Formation). Zootaxa, 2382, 1-71. Owen, R. (1861). Palaeontology, or a systematic summary of extinct animals and their geological relations (2nd ed., p. xvi+163). Adam and Black. Pacheco, C. P., Martinelli, A. G., Pavanatto, A. E. B., Soares, M. B., & Dias-Da-Silva, S. (2018). Prozostrodon brasiliensis, a probainognathian cynodont from the Late Triassic of Brazil: Second record and improvements on its dental anatomy. Historical Biology, 30, 475-485. Paes-Neto, V. D., Lacerda, M. B., Ezcurra, M. D., Raugust, T., Trotteyn, M., Soares, M. B., Schultz, C. L., Pretto, F. A., Francischini, H., & Martinelli, A. G. (2024). New rhadinosuchine proterochampsids from the late Middle-early Late Triassic of southern Brazil enhance the diversity of archosauriforms. The Anatomical Record, 307(4), 851-889. https://doi.org/10.1002/ar.25294. Peecook, B. R., Sidor, C. A., Nesbitt, S. J., Smith, R. M. H., Steyer, J. S., & Angielczyk, K. D. (2013). A new silesaurid from the upper Ntawere Formation of Zambia (Middle Triassic) demonstrates the rapid diversification of Silesauridae (Avemetatarsalia: Dinosauriformes). Journal of Vertebrate Paleontology, 33, 1127-1137. Perez Loinaze, V. S., Vera, E. I., Fiorelli, L. E., & Desojo, J. B. (2018). Palaeobotany and palynology of coprolites from the Late Triassic Chañares Formation of Argentina: Implications for vegetation provinces and the diet of dicynodonts. Palaeogeography, Palaeoclimatology, Palaeoecology, 502, 31-51. https://doi.org/10.1016/j.palaeo.2018.04.003. Pol, D., & Escapa, I. H. (2009). Unstable taxa in cladistic analysis: Identification and the assessment of relevant characters. Cladistics, 25(5), 515-527. Ponce, D., Trotteyn, M. J., Cerda, I., Fiorelli, L., & Desojo, J. B. (2021). Osteohistology and palaeobiological inferences of proterochampsids (Eucocropoda: Proterochampsia) from Chañares Formation (late Ladinian-Upper Triassic), La Rioja, Argentina. Journal of Vertebrate Paleontology, 41(2), e1926273. Reig, O. A. (1958). Primeros datos descriptivos sobre nuevos reptiles arcosaurios del Triásico de Ischigualasto (San Juan, Argentina). Revista de la Asociación Geológica Argentina, 13, 257-270. Reig, O. A. (1963). La presencia de dinosaurios saurísquios en los "Estratos de Ischigualasto" (Mesotriásico superior) de las provincias de San Juan y La Rioja (República Argentina). Ameghiniana, 3, 3-20. Rogers, R. R., Arcucci, A. B., Abdala, F., Sereno, P. C., Forster, C. A., & May, C. L. (2001). Paleoenvironment and taphonomy of the Chañares Formation tetrapod assemblage (Middle Triassic), northwestern Argentina: Spectacular preservation in volcanogenic concretions. PALAIOS, 16(5), 461-481. Romer, A. S. (1967). The Chañares (Argentina) Triassic reptile fauna. III. Two new gomphodonts, Massetognathus pascuali and M. teruggii. Breviora, 264, 1-25. Romer, A. S. (1969a). The Brazilian Triassic cynodont reptiles Belesodon and Chiniquodon. Breviora, 332, 1-16. Romer, A. S. (1969b). The Chañares (Argentina) Triassic reptile fauna. V: A new chiniquodontidae cynodont, Probelesodon lewisi. Breviora, 333, 1-24. Romer, A. S. (1970). The Chañares (Argentina) Triassic reptile fauna. VI. A chiniquodontid cynodont with incipient squamosal-dentary jaw articulation. Breviora, 344, 1-18. Romer, A. S. (1972). The Chañares (Argentina) Triassic reptile fauna. XVII. The Chañares Gomphodonts. Breviora, 396, 1-9. Romer, A. S. (1973a). The Chañares (Argentina) Triassic reptile fauna. XX. Summary. Breviora, 413, 1-20. Romer, A. S. (1973b). The Chanares (Argentina) Triassic reptile fauna. XVIII. Probelesodon minor, a new species of carnivorous cynodont; family Probainognathidae nov. Breviora, 401, 1-4. Romer, A. S., & Cox, C. B. (1962). The fossiliferous Triassic deposits of Ischigualasto, Argentina, and preliminary description of Ischigualastia, a new genus of dicynodont. Breviora, 156, 1-9. Romer, A. S., & Jensen, J. A. (1966). The Chañares (Argentina) Triassic reptile fauna. II. Sketch of the geology of the río Chañares-río Gualo region. Breviora, 252, 1-20. Romer, A. S., & Lewis, A. D. (1973). The Chañares (Argentina) Triassic reptile fauna. XIX. Postcranial materials of the cynodonts Probelesodon and Probainognathus. Breviora, 407, 1-26. Rougier, G. W., Martinelli, A. G., & Forasiepi, A. M. (2021). Mesozoic mammals from South America and their forerunners. Springer Earth System Sciences. Rowe, T. (1988). Definition, diagnosis and origin of Mammalia. Journal of Vertebrate Paleontology, 8, 241-264. Ruta, M., Botha-Brink, J., Mitchell, S. A., & Benton, M. J. (2013). The radiation of cynodonts and the ground plan of mammalian morphological diversity. Proceedings of the Royal Society B, 280, 20131865. Schmitt, M. R., Martinelli, A. G., Kaiuca, J. F. L., & Schultz, C. L. (2024). Old fossil findings in the Upper Triassic rocks of southern Brazil improve diversity of traversodontid cynodonts (Therapsida, Cynodontia). The Anatomical Record, 307(4), 1474-1514. https://doi.org/10.1002/ar.25244. Schmitt, M. R., Martinelli, A. G., Melo, T. P., & Soares, M. B. (2019). On the occurrence of the traversodontid Massetognathus ochagaviae (Synapsida, Cynodontia) in the early Late Triassic Santacruzodon Assemblage Zone (Santa Maria Supersequence, southern Brazil): Taxonomic and biostratigraphic implications. Jounal of South American Earth Sciences, 93, 36-50. Schultz, C. L., Martinelli, A. G., Soares, M. B., Pinheiro, F. L., Kerber, L., Horn, B. L. D., Pretto, F. A., Müller, R. T., & Melo, T. P. (2020). Triassic faunal successions of the Paraná Basin, southern Brazil. Jounal of South American Earth Sciences, 104, 102846. Sereno, P. C. (2013). Preface. Society of Vertebrate Paleontology memoir 12. Journal of Vertebrate Paleontology, 32(supplement to 6), 1-9. Sidor, C. A., & Smith, R. M. H. (2004). A new galesaurid (Therapsida: Cynodontia) from the Lower Triassic of South Africa. Palaeontology, 47, 535-556. Sidor C. A., & James A. Hopson J. A. (2017). Cricodon metabolus (Cynodontia: Gomphodontia) from the Triassic Ntawere Formation of northeastern Zambia: patterns of tooth replacement and a systematic review of the Trirachodontidae. Journal of Vertebrate Paleontology, 37(1, supplement), 39-64. https://doi.org/10.1080/02724634.2017.1410485. Sill, W. D. (1970). Scaphonyx sanjuanensis, nuevo rincosaurio (Reptilia) de la Formación Ischigualasto, Triásico de San Juan, Argentina. Ameghiniana, 7, 341-354. Soares, M. B., Martinelli, A. G., & Oliveira, T. V. (2014). A new prozostrodontian cynodont (Therapsida) from the Late Triassic Riograndia Assemblage one (Santa Maria Supersequence) of southern Brazil. Anais da Academia Brasileira de Ciências, 86, 1673-1691. Soares, M. B., Schultz, C. L., & Horn, B. L. D. (2011). New information on Riograndia guaibensis Bonaparte, Ferigolo & Ribeiro, 2001 (Eucynodontia, Tritheledontidae) from the Late Triassic of southern Brazil: Anatomical and biostratigraphic implications. Anais da Academia Brasileira de Ciências, 83, 329-354. Spalletti, L. A., Fanning, C. M., & Rapela, C. W. (2008). Dating the Triassic continental rift in the southern Andes: The Potrerillos Formation, Cuyo Basin, Argentina. Acta Geologica, 6, 267-283. https://doi.org/10.1344/105.000000256. Stefanello, M., Kerber, L., Martinelli, A. G., & Dias-Da-Silva, S. (2020). A new prozostrodontian cynodont (Eucynodontia, Probainognathia) from the Upper Triassic of southern Brazil. Journal of Vertebrate Paleontology, 40(3), e1782415. https://doi.org/10.1080/02724634.2020.1782415. Stefanello, M., Martinelli, A. G., Müller, R. T., Dias-da-Silva, S., & Kerber, L. (2023). A complete skull of a stem mammal from the Late Triassic of Brazil illuminates the early evolution of prozostrodontian cynodonts. Journal of Mammalian Evolution, 30, 299-317. https://doi.org/10.1007/s10914-022-09648-y. Sterli, J., Rougier, G., & de la Fuente, M. (2007). Anatomy and relationships of Palaeochersis talampayensis, a Late Triassic turtle from Argentina. Palaeontographica Abteilung A, 281(1), 1-61. Sues, H.-D. (1986). The skull and dentition of two tritylodontid synapsids from the Lower Jurassic of western North America. Bulletin of the Museum of Comparative Zoology, 151, 217-268. Sues, H.-D. (2016). Dating the origin of dinosaurs. Proceedings of the National Academy of Sciences of the United States of America, 113, 480-481. Szczygielski, T., & Sulej, T. (2023). Woznikella triradiata n. gen., n. sp., a new kannemeyeriiform dicynodont from the Late Triassic of northern Pangea and the global distribution of Triassic dicynodonts. Comptes Rendus Palevol, 22(16), 279-406. Velazco, P. M., Buczek, A. J., & Novacek, M. J. (2017). Two new tritylodontids (Synapsida, Cynodontia, Mammaliamorpha) from the Upper Jurassic, southwestern Mongolia. American Museum Novitates, 3874, 1-35. https://doi.org/10.1206/3874.1. von Baczko, M. B., Desojo, J. B., & Ponce, D. (2019). Postcranial anatomy and osteoderm histology of Riojasuchus tenuisceps and a phylogenetic update on Ornithosuchidae (Archosauria, Pseudosuchia). Journal of Vertebrate Paleontology, 39(5), e1693396. von Huene, F. (Ed.). (1936). Lieferung 2. Cynodontia. In Die fossilen reptilien des südamerikanischen gondwanalandes. Ergebnisse der Sauriergrabungen in Südbrasilien 1928/29 (pp. 83-160). C. H. Beck'sche Verlagsbuchhandlung. Wallace, R. V. S., Martínez, R., & Rowe, T. (2019). First record of a basal mammaliamorph from the early Late Triassic Ischigualasto Formation of Argentina. PLoS One, 14, e0218791. Wynd, B. M., Peecook, B. R., Whitney, M. R., & Sidor, C. A. (2017). The first occurrence of Cynognathus crateronotus (Cynodontia: Cynognathia) in Tanzania and Zambia, with implications for the age and biostratigraphic correlation of Triassic strata in southern Pangea. Journal of Vertebrate Paleontology, 37(sup1), 228-239. https://doi.org/10.1080/02724634.2017.1421548. Yeh, H. K. (1959). New dicynodont from Sinokannemeyeria-fauna from Shansi. Vertebrata Palasiatica, 3(4), 187-204.
معلومات مُعتمدة:	PICT 2020-SERIEA-01498 Agencia Nacional de Investigaciones Científicas y Técnicas; PICT 2018-1186 Agencia Nacional de Investigaciones Científicas y Técnicas; PICT 2018-0717 Agencia Nacional de Investigaciones Científicas y Técnicas; PICT 2014-0609 Agencia Nacional de Investigaciones Científicas y Técnicas; PICT 2012-0925 Agencia Nacional de Investigaciones Científicas y Técnicas; PICT 2010-0207 Agencia Nacional de Investigaciones Científicas y Técnicas; 28720210101137 PIBAA; Gobierno de La Rioja
فهرسة مساهمة:	Keywords: Chiniquodontidae; Cynodontia; Probainognathia; South America; taxonomy
تواريخ الأحداث:	Date Created: 20240129 Date Completed: 20240318 Latest Revision: 20240318
رمز التحديث:	20240318
DOI:	10.1002/ar.25388
PMID:	38282519
قاعدة البيانات:	MEDLINE

View record at Wiley

Full Text Finder

الوصف
تدمد:	1932-8494
DOI:	10.1002/ar.25388