A previous publication (Wilson & Angus 2005) gave information on the karyotypes of species of Oniticellini and Onthophagini studied by C. J. Wilson in her Ph. D. research (Wilson 2002). The present paper reports on the remaining Scarabaeidae (Gymnopleurus geoffroyi Illiger, Copris lunaris (Linnaeus) and Onitis belial Fabricius) included in that research, with additional material collected by R. B. Angus (European and Cypriot material) and D. J. Mann (Malaysian material).
