A geometrical formulation for adjoint-symmetries as one-forms is studied for general partial differential equations (PDEs), which provides a dual counterpart of the geometrical meaning of symmetries as tangent vector fields on the solution space of a PDE. Two applications of this formulation are presented. Additionally, for systems of evolution equations, adjoint-symmetries are shown to have another geometrical formulation given by one-forms that are invariant under the flow generated by the system on the solution space. This result is generalized to systems of evolution equations with spatial constraints, where adjoint-symmetry one-forms are shown to be invariant up to a functional multiplier of a normal one-form associated with the constraint equations. All of the results are applicable to the PDE systems of interest in applied mathematics and mathematical physics.